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Nature-based solutions (NBS) are a promising and innovative approach to address multiple sustainability challenges faced by cities. Yet, NBS are not integrated into mainstream urban development practices. Based on a qualitative comparative case study of Germany, Hungary, the Netherlands, Spain, Sweden, and the United Kingdom, this study shows how barriers to mainstreaming urban NBS are shaped by the structural conditions in urban infrastructure regimes, which offers an improved, context-sensitive understanding of why such barriers persist. We identify underlying structural conditions shaping seven key barriers to urban NBS: limited collaborative governance, knowledge, data and awareness challenges, low private sector engagement, competition over urban space, insufficient policy development, implementation and enforcement, insufficient public resources, and challenging citizen engagement. This …